Ph.D. Institue of Oceanology, Acad. Sci. USSR Moscow 1967
- Developmental biology of fishes, especially larval organogenesis. Reproductive physiology concerning applied research on sexual maturation in broodstocks of cultured fish. Environmental effects on early life stages of fish, including bioassay designs for toxicological research with embryos, larvae, and juveniles.
Areas of Emphasis
- Physiological Ecology, Conservation Ecology, Integrative Ecology
- Animal Science